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Granulocyte-macrophage colony-stimulating factor (GM-CSF), also known as colony-stimulating factor 2 (CSF2), is a monomeric glycoprotein secreted by macrophages, T cells, mast cells, natural killer cells, endothelial cells and fibroblasts, which function as a cytokine. Unlike granulocyte colony-stimulating factor, which specifically promotes the proliferation and maturation of neutrophils, GM-CSF affects many more cell types, especially macrophages and eosinophils.