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MUC4 membrane-bound mucin is a large O glycoprotein involved in epithelial homeostasis. On the surface of cancer cells, MUC4 interacts with ErbB2 receptors through the EGF domain to promote cell proliferation and migration. MUC4 is highly regarded as a therapeutic target in pancreatic cancer as it is not expressed in healthy pancreas, while it is neoexpressed in early preneoplastic stages (PanINs).